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Hello all!

Is there a way to communicate between a atmega328 and the Z-UNO? The TX&RX pin on the mega is already in use. maybe possible to use I2c?

You can use SPI, I2C or even own digital protocol on a pin (or on many pins). This is up to your imagination

You could use SoftwareSerial on mega328 too, but I prefer I2C, because you could use only 2 pins for a number of devices. In that case you have to configure mega328 as I2C slave device.